[1] Korea Inst Ocean Sci & Technol, East Sea Environm Res Div, Uljin Gun, Gyeongbuk, South Korea
[2] Korea Maritime & Ocean Univ, Ocean Sci & Technol Sch, Dept Convergence Study Ocean Sci & Technol, Busan, South Korea
[3] Korea Univ Sci & Technol, Dept Marine Biotechnol, Daejeon, South Korea
[4] Korea Inst Ocean Sci & Technol, Marine Ecosyst Res Div, 787 Haean Ro, Ansan 426744, South Korea
[5] Korea Univ Sci & Technol, Dept Marine Biol, Daejeon, South Korea