[1] Korea Univ Sci & Technol, Dept Marine Biol, Daejeon, South Korea
[2] Korea Inst Ocean Sci & Technol, Biol Oceanog & Marine Biol Div, Ansan, South Korea
[3] East Sea Fisheries Res Inst, Fisheries Resources & Environm Div, Kangnung, Gangwon Do, South Korea
[4] Rural Dev Adm, Natl Inst Anim Sci, Cheonan Si, Chungnam, South Korea
[5] Korea Inst Ocean Sci & Technol, East Sea Res Inst, Uljin Gun, Gyeongbuk, South Korea
[6] Korea Univ Sci & Technol, Dept Marine Biotechnol, Daejeon, South Korea
[7] Korea Maritime & Ocean Univ, Ocean Sci & Technol Sch, Dept Convergence Study Ocean Sci & Technol, Busan, South Korea