[1] Univ Illinois, Dept Cell & Dev Biol, Urbana, IL 61820 USA
[2] Univ Illinois, Inst Genom Biol, Urbana, IL 61820 USA
[3] Harvard Univ, Museum Comparat Zool, Cambridge, MA 02138 USA
[4] Harvard Univ, Dept Organism & Evolutionary Biol, Cambridge, MA 02138 USA