The fault-bounded Proterozoic metamorphic terranes lying to the E of the Tanzanian craton make up the Usagara tectonic domain and are a part of the transcontinental Mozambique Orogenic Belt (MB). The lithotectonic units in the MB of East Africa consist of comparable rock assemblages which underwent the same complex deformational history and are thought to represent large thrust sheets or nappes. Their shelf- and fore-deep terranes border the Tanzanian craton and make up the foreland terranes of the Pan-African Mozambique Belt. Granulite-gneiss nappes are ubiquitous in the orogen. Granulite-facies metamorphism, associated with recumbent folds, was due to crustal thickening, which took place during the collision between Gondwana fragments. Isotope data suggest a collision (and concomitant granulite-facies metamorphism) age of between 700 and 550 Ma. The orientations of planar and linear fabrics in the granulite-facies rocks of the Uluguru mountains are used to infer the relative crustal block motions during this collisional event. This Pan-African collisional event was characterized by NW-directed movements, oblique to the N-S trend of the orogen, and involved SE-directed backthrusting. The Ubendian Belt of Tanzania and the Aswa Shear Zone in Uganda and Kenya, which both bifurcate around the Tanzania craton, accommodated the tectonically thickened crust, created by the collisional event, through NW-SE sinistral strike-slip movements.