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Natl Agr & Food Res Org NARO, Inst Livestock & Grassland Sci, Div Meat Anim & Poultry Res, Meat Anim Biosyst Grp, 2 Ikenodai, Tsukuba, Ibaraki 3050901, Japan Natl Agr & Food Res Org NARO, Inst Livestock & Grassland Sci, Div Meat Anim & Poultry Res, Meat Anim Biosyst Grp, 2 Ikenodai, Tsukuba, Ibaraki 3050901, Japan