[1] Univ Liverpool, Inst Ageing & Chron Dis, Dept Eye & Vis Sci, Liverpool L7 8TX, Merseyside, England
[2] Univ Newcastle, Int Ctr Life, Inst Genet Med, Newcastle NE1 3BZ, England
[3] Spheritech Ltd, Heath Business & Tech Pk, Runcorn WA7 4QX, Cheshire, England