[1] UCL Inst Ophthalmol, Cell Biol, London, England
[2] Univ Sydney, Save Sight Inst, Sydney, NSW, Australia
[3] Univ Melbourne, Ctr Eye Res Australia, Royal Victorian Eye & Ear Hosp, Melbourne, Vic, Australia
[4] Univ Utah, Dept Ophthalmol & Visual Sci, John A Moran Eye Ctr, Salt Lake City, UT 84112 USA