[1] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Ciencias Biosistemat & Manejo Recursos Nat & Agr, Camino Ing Ramon Padilla Sanchez 2100, Zapopan 45200, Jalisco, Mexico
[2] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Dept Bot & Zool, Herbaria Luz Maria Villarreal Puga,Inst Bot, Camino Ing Ramon Padilla Sanchez 2100, Zapopan 45200, Jalisco, Mexico
[3] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Conacyt, Lab Nacl Identificac & Caracterizac Vegetal, Camino Ing Ramon Padilla 2100, Zapopan 45200, Jalisco, Mexico
[4] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Catedras Conacyt, Camino Ing Raman Padilla 2100, Zapopan 45200, Jalisco, Mexico