[1] Swedish Museum Nat Hist, Dept Phanerogam Bot, SE-10405 Stockholm, Sweden
[2] Uppsala Univ, Dept Systemat Bot, SE-75236 Uppsala, Sweden
[3] Royal Swedish Acad Sci, Bergius Fdn, SE-10405 Stockholm, Sweden
[4] Bergius Fdn, Dept Bot, SE-10691 Stockholm, Sweden
[5] Museum Natl Hist Nat, Dept Systemat & Evolut, Herbier Natl, CNRS,UMS 2700,USM 0602, F-75231 Paris, France