[1] Russian Acad Sci, Zool Inst, Univ Skaya Nab 1, St Petersburg 199034, Russia
[2] Tomsk State Univ, Lab Mesozo & Cenozo Continental Ecosyst, Prospekt Lenina 36, Tomsk 634050, Russia
[3] Russian Acad Sci, Borissiak Paleontol Inst, Profsoyuznaya 123, Moscow 117997, Russia
[4] St Petersburg State Univ, Dept Vertebrate Zool, Univ Skaya Nab 7-9, St Petersburg 199034, Russia