[1] Ocean Univ China, Inst Coastal Environm Pollut Control, Coll Environm Sci & Engi neering, Minist Educ Key Lab Marine Environm & Ecol, Qingdao 266100, Peoples R China
[2] Minist Nat Resources MNR, Inst Oceanog 1, Marine Bioresource & Environm Res Ctr, Key Lab Marine Ecoenvironm Sci & Technol, Qingdao 266061, Peoples R China
[3] Lab Marine Drugs & Bioprod, Pilot Natl Lab Marine Sci & Technol, Qingdao 266071, Peoples R China
[4] Shandong Univ, Inst Marine Sci & Technol, Qingdao 266200, Peoples R China
[5] Minist Nat Resources, Inst Oceanog 1, Key Lab Marine Sci & Numer Modeling, Qingdao 266061, Peoples R China