[1] Univ Fed Rural Amazonia, Museu Paraense Emilio Goeldi, Programa Posgrad Ciencias Biol Bot Trop, Ave Perimetral 1901, BR-66077830 Belem, Para, Brazil
[2] Univ Fed Pernambuco, Lab Biol Floral & Reprod, Ctr Biociencias, Ave Prof Moraes Rego 1235,Cidade Univ, BR-50670901 Recife, PE, Brazil
[3] Univ Gibraltar, Gibraltar Bot Gardens Campus,Red Sands Rd,POB 843, The Alameda GX11 1AA, Gibraltar
[4] Univ Brasilia, Inst Ciencias Biol, Programa Posgrad Bot, Caixa Postal 04457, BR-70910970 Brasilia, DF, Brazil