[1] Vietnam Acad Sci & Technol, Inst Ecol & Biol Resources, Dept Bot, 18 Hoang Quoc Viet, Hanoi 10072, Vietnam
[2] Grad Univ Sci & Technol, Vietnam Acad Sci & Technol, 18 Hoang Quoc Viet, Hanoi 10072, Vietnam
[3] Chinese Acad Sci, Key Lab Plant Resources Conservat & Sustainable U, South China Bot Garden, Guangzhou 510650, Peoples R China
[4] Chinese Acad Sci, Southeast Asia Biodivers Res Inst, Yezin 05282, Nay Pyi Taw, Myanmar
[5] Agharkar Res Inst, Biodivers & Palaeobiol Grp, GG Agarkar Rd, Pune 411004, Maharashtra, India
[6] Chungnam Natl Univ, Daejeon 34134, South Korea