[1] Univ Toronto, Dept Biol, Mississauga, ON L5L 1C6, Canada
[2] Univ Iowa, Coll Med, Howard Hughes Med Inst, Iowa City, IA 52242 USA
[3] Wageningen Univ, Entomol Lab, NL-6709 PD Wageningen, Netherlands
[4] KNAW, NIOO, Netherlands Inst Ecol, NL-3600 BG Maarssen, Netherlands
[5] Univ Illinois, Dept Entomol, Urbana, IL 61801 USA
[6] Univ Illinois, Neurosci Program, Urbana, IL 61801 USA