[1] Chinese Acad Sci, Key Lab Vertebrate Evolut & Human Origin, Inst Vertebrate Paleontol & Paleoanthropol, Beijing 100044, Peoples R China
[2] CAS Ctr Excellence Life & Paleoenvironment, Beijing 100044, Peoples R China
[3] Chinese Acad Sci, Inst Geol & Geophys, State Key Lab Lithospher Evolut, Beijing 100029, Peoples R China
[4] Univ Calif Davis, Dept Earth & Planetary Sci, Davis, CA 95616 USA