[1] Univ Aberdeen, Inst Med Sci, Immun Infect & Inflammat Programme, Aberdeen, Scotland
[2] Univ Cape Town, ICGEB, IIDMM, ZA-7925 Cape Town, South Africa
[3] Univ Ghent, Dept Resp Dis, Lab Immunoregulat & Mucosal Immunol, B-9000 Ghent, Belgium
[4] Univ Aberdeen, Dept Pathol, Inst Med Sci, Aberdeen, Scotland
[5] Univ Pittsburgh, Sch Med, Dept Pediat,Inst Pediat Res, Childrens Hosp Pittsburgh,Richard King Mellon Fdn, Pittsburgh, PA 15261 USA