[1] Inst Bot Darwinion, Buenos Aires, DF, Argentina
[2] Univ Nacl Agraria La Molina, Fac Ciencias, Herbario Dept Biol, Lima 12, Peru
[3] Univ Mayor San Andres, Inst Ecol, Herbario Nacl Bolivia, La Paz 10077, Bolivia
[4] Univ Buenos Aires, Fac Agron, Catedra Bot Gen, Buenos Aires, DF, Argentina