[1] Univ Hawaii Manoa, Int Pacific Res Ctr, Honolulu, HI 96822 USA
[2] Univ Hawaii Manoa, Dept Oceanog, Honolulu, HI 96822 USA
[3] Japan Agcy Marine Earth Sci & Technol, Yokohama, Kanagawa, Japan
[4] Univ Tokyo, Res Ctr Adv Sci & Technol, Tokyo, Japan