[1] US Geol Survey, Southwest Biol Sci Ctr, Tucson, AZ 85721 USA
[2] Univ Arizona, Biosyst Engn, Tucson, AZ 85721 USA
[3] Univ Technol Sydney, Fac Sci, Sch Life Sci, Sydney, NSW 2007, Australia
[4] AREEO, Soil Conservat & Watershed Management Res Dept, Isfahan Agr & Nat Resources Res & Educ Ctr, Esfahan 8174835117, Iran
[5] Univ Gottingen, Div Agron, Von Siebold Str 8, D-37075 Gottingen, Germany