[1] Univ La Reunion, UMR C53 Cirad Univ, Peuplements Vegetaux & Bioaggresseurs Milieu Trop, St Denis Messag 9, La Reunion, France
[2] Univ Antananarivo, Fac Sci Ment Biol & Ecol Vegetales, Antananarivo, Madagascar
[3] Swedish Museum Nat Hist, Dept Bot, Box 50007, SE-10405 Stockholm, Sweden
[4] Univ KwaZulu Natal, Sch Life Sci, Ctr Funct Biodivers, Bews Herbarium, Private Bag X01, ZA-3209 Scottsville, South Africa
[5] Univ Mauritius, Fac Sci, Trop Isl Biodivers Ecol & Conservat Pole Res, Reduit 80837, Mauritius
[6] Univ Antilles, EPHE, Sorbonne Univ, CNRS,Museum Natl Hist Nat,Inst Systemat Evolut Bi, CP 39,57 Rue Cuvier, F-75005 Paris, France
[7] Minist Agroind & Food Secur, Agr Serv, Mauritius Herbarium, RE Vaughan Bldg, Reduit 80835, Mauritius