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Univ Franche Comte, CHRU Besancon, COMUE Bourgogne Franche Comte Univ, Dept Virol,EPILAB Pathogens & Inflammat Lab,EA426, Besancon, France Univ Franche Comte, CHRU Besancon, COMUE Bourgogne Franche Comte Univ, Dept Virol,EPILAB Pathogens & Inflammat Lab,EA426, Besancon, France