[1] Univ Porto, Fac Ciencias, REQUIMTE, Dept Quim, P-4167007 Oporto, Portugal
[2] CIBIO, UP, Ctr Invest Biodiversidade & Recursos Genet, P-4485661 Vairao, Portugal
[3] Univ Porto, Fac Ciencias, Dept Zool Anthropol, P-4099002 Oporto, Portugal
[4] Univ Porto, Fac Ciencias, REQUIMTE, Dept Quim, P-4169007 Oporto, Portugal
[5] INRA, Lab Genet Poissons, F-78352 Jouy En Josas, France
[6] Univ Guelph, Dept Integrat Biol, Guelph, ON N1G 2W1, Canada