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Chinese Acad Agr Sci, Key Lab Anim Virol, Minist Agr, State Key Lab Vet Etiol Biol,Lanzhou Vet Res Inst, Lanzhou 730046, Gansu, Peoples R China Chinese Acad Agr Sci, Key Lab Anim Virol, Minist Agr, State Key Lab Vet Etiol Biol,Lanzhou Vet Res Inst, Lanzhou 730046, Gansu, Peoples R China