[1] Univ Guelph, Dept Integrat Biol, Guelph, ON N1G 2W1, Canada
[2] Univ Guelph, Biodivers Inst Ontario, Guelph, ON N1G 2W1, Canada
[3] Canadian Natl Collect Insects, Ottawa, ON K1A 0C6, Canada
[4] Univ Illinois, Dept Entomol, Urbana, IL 61801 USA
[5] Univ Liege, Gembloux Agrobio Tech, Unite Entomol Fonct & Evolut, B-1030 Gembloux, Belgium
[6] IRSNB, Dept Entomol, B-1000 Brussels, Belgium
[7] Univ Penn, Dept Biol, Philadelphia, PA 19104 USA