[1] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol, Beijing, Peoples R China
[2] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Key Lab Vertebrate Evolut & Human Origins, Beijing, Peoples R China
[3] CAS Ctr Excellence Life & Paleoenvironm, Beijing, Peoples R China
[4] Ctr Nacl Invest Evoluc Humana CENIEH, Burgos, Spain
[5] Univ Chinese Acad Sci, Beijing, Peoples R China
[6] Univ Burgos, Doctorado Interuniv Evoluc Humana Paleoecol Cuate, Burgos, Spain