[1] Calif State Univ Channel Isl, Camarillo, CA 93012 USA
[2] Geol Survey Canada Atlantic, Nat Resources Canada, Bedford Inst Oceanog, Dartmouth, NS B2Y 4A2, Canada
[3] Fisheries & Oceans Canada, Sci Branch, Maritimes Reg, Biol Stn, St Andrews, NB E5B 2L9, Canada
[4] NOAA Fisheries, US Dept Commerce, NE Fisheries Sci Ctr, James J Howard Marine Sci Lab, Sandy Hook, NJ 07732 USA