[1] Univ Kansas, Amer Museum Nat Hist, Div Invertebrate Zool Entomol, Lawrence, KS 66045 USA
[2] Univ Kansas, Nat Hist Museum, Div Entomol, Lawrence, KS 66045 USA
[3] Univ Kansas, Dept Ecol & Evolutionary Biol, Lawrence, KS 66045 USA
[4] King Saud Univ, Dept Plant Protect, Coll Food & Agr Sci, Riyadh, Saudi Arabia
[5] Suez Canal Univ, Dept Plant Protect, Fac Agr, Ismailia, Egypt