[1] Chinese Acad Sci, Xishuangbanna Trop Bot Garden, Ctr Integrat Conservat, Plant Phylogenet & Conservat Grp, Kunming, Yunnan, Peoples R China
[2] Univ Chinese Acad Sci, Beijing, Peoples R China
[3] Chinese Acad Sci, Xishuangbanna Trop Bot Garden, Herbarium HITBC, Menglun, Yunnan, Peoples R China
[4] Univ Hamburg, Biozentrum Klein Flottbek, Hamburg, Germany
[5] Chinese Acad Sci, Kunming Inst Bot, Herbarium KUN, Kunming, Yunnan, Peoples R China