[1] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China
[2] USDA ARS, Systemat Entomol Lab, Washington, DC 20250 USA
[3] Univ Chinese Acad Sci, Beijing 100039, Peoples R China
[4] Kerala Agr Univ, Dept Entomol, Trivandrum 695522, Kerala, India