[1] Finnish Meteorol Inst, POB 503, Helsinki 00101, Finland
[2] Univ Helsinki, Dept Geosci & Geog, Gustaf Hallstromin Katu 2a,POB 64, Helsinki 00014, Finland
[3] Swedish Univ Agr Sci, Dept Forest Ecol & Management, PO 901 83, Umea, Sweden
[4] Univ Helsinki, Dept Forest Sci, POB 27, Helsinki 00014, Finland
[5] Norwegian Meteorol Inst, Blindern,POB 43, N-0313 Oslo, Norway