[1] Queens Univ Belfast, Ctr Infect & Immun, Belfast, Antrim, North Ireland
[2] Univ Leicester, Inst Lung Hlth, Leicester, Leics, England
[3] Royal Brompton & Harefield NHS Fdn Trust, NIHR Resp Dis, Biomed Res Unit, London, England
[4] Univ Manchester, NW Lung Ctr, Manchester, Lancs, England
[5] Univ London Imperial Coll Sci Technol & Med, London, England