[1] South African Natl Biodivers Inst, Compton Herbarium, Private Bag X7, ZA-7735 Cape Town, South Africa
[2] Univ KwaZulu Natal, Res Ctr Plant Growth & Dev, Sch Biol & Conservat Sci, Private Bag X01, ZA-3209 Pietermaritzburg, Scottsville, South Africa
[3] Missouri Bot Garden, BA Krukoff Curator African Bot, POB 299, St Louis, MO 63166 USA
[4] Threatened Species Programme, Durban, South Africa