[1] Univ Alcala de Henares, Dept Ciencias Vida, UD Ecol, E-28871 Alcala De Henares, Spain
[2] Univ Rey Juan Carlos, Dept Biol & Geol, Area Biodiversidad & Conservac, E-28933 Mostoles, Madrid, Spain
[3] Karlsruhe Inst Technol, Inst Meteorol & Climate Res Atmospher Environm Re, Garmisch Partenkirchen, Germany
[4] Univ Nacl Colombia, Fac Ciencias, Dept Biol, Lab Ecol Paisaje & Modelac Ecosistemas, Bogota, Colombia