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Chinese Acad Agr Sci, Inst Grassland Res, Key Lab Biohazard Monitoring & Green Prevent & Con, Minist Agr & Rural Affairs, Hohhot 010010, Inner Mongolia, Peoples R China Chinese Acad Agr Sci, Inst Grassland Res, Key Lab Biohazard Monitoring & Green Prevent & Con, Minist Agr & Rural Affairs, Hohhot 010010, Inner Mongolia, Peoples R China