[1] Univ Aberdeen, Sch Biol Sci, Aberdeen, Scotland
[2] Columbia Univ, Dept Biol Sci, New York, NY 10027 USA
[3] Uppsala Univ, Dept Ecol & Genet, Uppsala, Sweden
[4] Univ Zurich, Dept Evolutionary Biol & Environm Studies, Zurich, Switzerland
[5] Univ Oslo, Dept Biosci, Ctr Ecol & Evolutionary Synth CEES, Oslo, Norway
[6] New Zealand Inst Plant & Food Res Ltd, Nelson, New Zealand
[7] Univ Auckland, Sch Biol Sci, Auckland, New Zealand