[1] Univ Southampton, Ctr Dev Origins Hlth & Dis, Southampton SO16 5YA, Hants, England
[2] Southampton Gen Hosp, Wessex Cardiothorac Unit, Southampton SO16 6YD, Hants, England
[3] Univ London Royal Vet Coll, Dept Vet Reprod, London AL9 7TA, England
[4] GKT Hosp, Div Reprod Hlth Endorcrinol & Dev, London SE1 7EH, England