[1] Ehime Univ, Ctr Marine Environm Studies, Matsuyama, Ehime, Japan
[2] Minist Nat Resources, Inst Oceanog 2, State Key Lab Satellite Ocean Environm Dynam, Hangzhou, Zhejiang, Peoples R China
[3] IPB Univ, Fac Fisheries & Marine Sci, Dept Aquat Resources Management, Bogor, Indonesia
[4] IPB Univ, Ctr Coastal & Marine Resources Studies, Bogor, Indonesia
[5] Nagasaki Univ, Grad Sch Fisheries & Environm Sci, Nagasaki, Japan