[1] Penn State Univ, Dept Anthropol, University Pk, PA 16802 USA
[2] Univ Calif San Diego, Dept Pediat, San Diego, CA 92161 USA
[3] Univ Hawaii, John A Burns Sch Med, Dept Anat Biochem & Physiol, Honolulu, HI 96813 USA
[4] Stowers Inst Med Res, Kansas City, MO USA
[5] Univ Kansas, Med Ctr, Dept Anat & Cell Biol, Kansas City, KS 66160 USA