[1] Univ San Francisco Quito USFQ, Inst iBIOTROP, Museo Zool, Quito 170901, Ecuador
[2] Univ San Francisco Quito USFQ, Inst iBIOTROP, Lab Zool Terrestre, Quito 170901, Ecuador
[3] Univ San Francisco Quito USFO, Colegio Ciencias Biol & Ambientales COCIBA, Quito, Ecuador
[4] Inst Nacl Biodiversidad, Unidad Invest, Quito, Ecuador
[5] Univ Porto, Ctr Invest Biodiversidade & Recursos Genet, CIBIO InBIO, P-4485661 Vairao, Portugal
[6] Pontificia Univ Catolica Ecuador, Escuela Biol, Museo Zool, Quito, Ecuador
[7] Real Jardin Bot RJB CSIC, Plaza Murillo 2, Madrid 28014, Spain