[1] Univ Western Australia, UWA Oceans Inst, Crawley, WA 6009, Australia
[2] Univ Western Australia, Sch Biol Sci, Crawley, WA 6009, Australia
[3] Inst Marine Res, His, Bergen, Norway
[4] Bedford Inst Oceanog, Fisheries & Oceans Canada, Dartmouth, NS, Canada
[5] Roskilde Univ, Dept Sci & Environm, Univ Vej 1, DK-4000 Roskilde, Denmark
[6] Newcastle Univ, Sch Nat & Environm Sci, Newcastle Upon Tyne NE1 7RU, Tyne & Wear, England