[1] Wageningen Univ, Anim Breeding & Genom Ctr, NL-6700 AH Wageningen, Netherlands
[2] Wageningen Univ, Behav Ecol Grp, NL-6700 AH Wageningen, Netherlands
[3] Wageningen UR, Livestock Res, NL-8200 AB Lelystad, Netherlands
[4] Wageningen Univ, Adaptat Physiol Grp, NL-6700 AH Wageningen, Netherlands
[5] Univ Utrecht, UIPS, Dept Psychopharmacol, NL-3508 TB Utrecht, Netherlands
[6] Univ Utrecht, Rudolf Magnus Inst Neurosci, NL-3508 TB Utrecht, Netherlands
[7] Univ Groningen, NL-9747 AG Groningen, Netherlands
[8] Univ Utrecht, Fac Vet Med, Dept Farm Anim Hlth, Emot & Cognit Grp, NL-3584 CL Utrecht, Netherlands
[9] Univ Utrecht, Rudolf Magnus Inst Neurosci, NL-3584 CL Utrecht, Netherlands
[10] Louis Bolk Inst, NL-3972 LA Driebergen, Netherlands
[11] Univ Bristol, Sch Vet Sci, Bristol BS40 5DU, Avon, England