[1] Nat Hist Museum, Terreaux 14, CH-2000 Neuchatel, Switzerland
[2] Agr Res Council, Private Bag X134, ZA-0121 Queenswood, South Africa
[3] Univ KwaZulu Natal, Sch Biol & Conservat Sci, Private Bag X01, ZA-3209 Pietermaritzburg, South Africa
[4] Univ Kiel, Zool Museum, Hegewischstr 3, D-24105 Kiel, Germany
[5] Nat Hist Museum, Dept Life Sci, Cromwell Rd, London SW7 5BD, England