[1] Pontificia Univ Catolica Valparaiso, Sch Marine Sci, Valparaiso, Chile
[2] UiT Arctic Univ Norway, Fac Biosci Fisheries & Econ, Tromso, Norway
[3] Leibniz Ctr Trop Marine Res, Bremen, Germany
[4] Univ Bremen, Fac Biol Chem, Bremen, Germany
[5] GEOMAR Helmholz Ctr Ocean Res Kiel, Kiel, Germany