[1] Nature Conservancy, Caribbean Program, Coral Gables, FL 33134 USA
[2] Gulf & Caribbean Fisheries Inst, Marathon, FL 33050 USA
[3] World Conservat Monitoring Ctr, United Nations Environm Programme, Cambridge CB3 0DL, England
[4] Univ So Mississippi, Dept Geog & Geol, Hattiesburg, MS 39406 USA