[1] Univ Tubingen, Senckenberg Ctr Human Evolut & Palaeoenvironm HEP, Paleoanthropol, Tubingen, Germany
[2] Univ Oslo, Dept Biosci, CEES, Oslo, Norway
[3] George Washington Univ, Dept Anthropol, Ctr Adv Study Human Paleobiol, Washington, DC 20052 USA
[4] Chatham Univ, Dept Biol, Pittsburgh, PA USA
[5] Assoc Vahatra, Antananarivo, Madagascar
[6] SUNY Stony Brook, Dept Anat Sci, Stony Brook, NY 11794 USA