[1] Univ Guelph, Ctr Genet Improvement Livestock, Guelph, ON, Canada
[2] Qualitas AG, Zug, Switzerland
[3] Bern Univ Appl Sci, Sch Agr Forest & Food Sci, Zollikofen, Switzerland
[4] Univ Guelph, Dept Anim Biosci, Ctr Genet Improvement Livestock, Guelph, ON, Canada
[5] Swiss Fed Inst Technol, Inst Agr Sci, Unit Anim Genet, Zurich, Switzerland
[6] Res Inst Organ Agr FiBL, Dept Anim Sci, Frick, Switzerland