[1] Univ London Imperial Coll Sci Technol & Med, Dept Biol Sci, Ascot SL5 7PY, Berks, England
[2] Nat Hist Museum, Dept Entomol, London SW7 5BD, England
[3] Univ Guelph, Biodivers Inst Ontario, Guelph, ON N1G 2W1, Canada
[4] Czech Acad Sci, Fac Sci, Univ South Bohemia, Ceske Budejovice 37005, Czech Republic
[5] Czech Acad Sci, Ctr Biol, Ceske Budejovice 37005, Czech Republic
[6] Chulalongkorn Univ, Fac Sci, Dept Biol, Bkk 10330, Thailand