[1] South African Natl Biodivers Inst, Compton Herbarium, ZA-7735 Claremont, South Africa
[2] Univ KwaZulu Natal, Sch Life Sci, Res Ctr Plant Growth & Dev, ZA-3209 Scottsville, South Africa
[3] Missouri Bot Garden, BA Krukoff Curator African Bot, St Louis, MO 63166 USA
[4] Geo Potts Herbarium, ZA-9300 Bloemfontein, South Africa