[1] Univ Cambridge, Dept Clin Neurosci, Cambridge CB2 0XY, England
[2] Univ Toronto, Ctr Res Neurodegenerat Dis, Toronto, ON, Canada
[3] Univ Toronto, Toronto Western Hosp, Div Neurol, Dept Med, Toronto, ON M5T 2S8, Canada
[4] Washington Univ, Sch Med, Dept Neurol, St Louis, MO 63110 USA