[1] MNHN, Dept Systemat & Evolut, CNRS, UMR7205, F-75231 Paris 05, France
[2] Univ Cocody Abidjan, UFR Biosci, Abidjan, Cote Ivoire
[3] Univ Douala, Fac Sci, Dept Anim Biol Organisms, Douala, Cameroon
[4] Univ Swaziland, Dept Biol Sci, Kwaluseni, Eswatini
[5] Univ Pretoria, Mammal Res Inst, Dept Zool & Entomol, ZA-0002 Pretoria, South Africa
[6] Severtsov Inst Ecol & Evolut, Moscow, Russia